Leucine and arginine enhance milk fat and milk protein synthesis via the CaSR/Gi/mTORC1 and CaSR/Gq/mTORC1 pathways.

Background and aims: Amino acids (AAs) not only constitute milk protein but also stimulate milk synthesis through the activation of mTORC1 signaling, but which amino acids that have the greatest impact on milk fat and protein synthesis is still very limited. In this study, we aimed to identify the m...

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Publicado en:European Journal of Nutrition Vol. 62; no. 7; pp. 2873 - 2891
Autores principales: Li, Qihui, Chen, Jiaming, Liu, Jiaxin, Lin, Tongbin, Liu, Xinghong, Zhang, Shuchang, Yue, Xianhuai, Zhang, Xiaoli, Zeng, Xiangfang, Ren, Man, Guan, Wutai, Zhang, Shihai
Formato: pictorial research tables/charts Journal Article
Publicado: Springer Nature Oct2023
Acceso en línea:Ver este registro en EBSCOhost
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      dt: Oct2023
      vid: 62
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      pub: Springer Nature
      place: New York, New York
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        10.1007/s00394-023-03197-7
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        atl: Leucine and arginine enhance milk fat and milk protein synthesis via the CaSR/Gi/mTORC1 and CaSR/Gq/mTORC1 pathways.
      aug:
        au:
          Li, Qihui
          Chen, Jiaming
          Liu, Jiaxin
          Lin, Tongbin
          Liu, Xinghong
          Zhang, Shuchang
          Yue, Xianhuai
          Zhang, Xiaoli
          Zeng, Xiangfang
          Ren, Man
          Guan, Wutai
          Zhang, Shihai
        affil: Guangdong Province Key Laboratory of Animal Nutrition Control, College of Animal Science, South China Agricultural University, 510642, Guangzhou, China
      sug:
        subj:
          Leucine Pharmacodynamics
          Arginine Pharmacodynamics
          Dietary Fats Metabolism
          Milk Proteins Metabolism
          Receptors, G-Protein-Coupled Drug Effects
          Intracellular Signaling Peptides and Proteins Drug Effects
          Membrane Proteins Drug Effects
          Calcium Binding Proteins Drug Effects
          Signal Transduction Drug Effects
          In Vitro Studies
          Animal Studies
          Mice
          Swine
          Epithelial Cells Drug Effects
          Cell Line Drug Effects
          Epithelial Cells Metabolism
          Cell Line Metabolism
          Lactation
          Gene Expression Drug Effects
          Fatty Acid-Binding Proteins Drug Effects
          Acyltransferases Drug Effects
          Transcription Factors Drug Effects
          Caseins Drug Effects
          Polymerase Chain Reaction
          Blotting, Western
          Descriptive Statistics
          Funding Source
      ab: Background and aims: Amino acids (AAs) not only constitute milk protein but also stimulate milk synthesis through the activation of mTORC1 signaling, but which amino acids that have the greatest impact on milk fat and protein synthesis is still very limited. In this study, we aimed to identify the most critical AAs involved in the regulation of milk synthesis and clarify how these AAs regulate milk synthesis through the G-protein-coupled receptors (GPCRs) signaling pathway. Methods: In this study, a mouse mammary epithelial cell line (HC11) and porcine mammary epithelial cells (PMECs) were selected as study subjects. After treatment with different AAs, the amount of milk protein and milk fat synthesis were detected. Activation of mTORC1 and GPCRs signaling induced by AAs was also investigated. Results: In this study, we demonstrate that essential amino acids (EAAs) are crucial to promote lactation by increasing the expression of genes and proteins related to milk synthesis, such as ACACA, FABP4, DGAT1, SREBP1, α-casein, β-casein, and WAP in HC11 cells and PMECs. In addition to activating mTORC1, EAAs uniquely regulate the expression of calcium-sensing receptor (CaSR) among all amino-acid-responsive GPCRs, which indicates a potential link between CaSR and the mTORC1 pathway in mammary gland epithelial cells. Compared with other EAAs, leucine and arginine had the greatest capacity to trigger GPCRs (p-ERK) and mTORC1 (p-S6K1) signaling in HC11 cells. In addition, CaSR and its downstream G proteins Gi, Gq, and Gβγ are involved in the regulation of leucine- and arginine-induced milk synthesis and mTORC1 activation. Taken together, our data suggest that leucine and arginine can efficiently trigger milk synthesis through the CaSR/Gi/mTORC1 and CaSR/Gq/mTORC1 pathways. Conclusion: We found that the G-protein-coupled receptor CaSR is an important amino acid sensor in mammary epithelial cells. Leucine and arginine promote milk synthesis partially through the CaSR/Gi/mTORC1 and CaSR/Gq/mTORC1 signaling systems in mammary gland epithelial cells. Although this mechanism needs further verification, it is foreseeable that this mechanism may provide new insights into the regulation of milk synthesis.
      pubtype: Academic Journal
      doctype:
        pictorial
        research
        tables/charts
        Journal Article
      ougenre: Article
    language: English
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