The Structures and Electronic Configuration of Compound I Intermediates of Helicobacter pylori and Penicillium vitale Catalases Determined by X-ray Crystallography and QM/MM Density Functional Theory Calculations.

The structures of Helicobacter pylori (HPC) and Penicillium vitale (PVC) catalases, each with two subunits in the crystal asymmetric unit, oxidized with peroxoacetic acid are reported at 1.8 and 1.7 Å resolution, respectively. Despite the similar oxidation conditions employed, the iron-oxygen coordi...

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Publicado en:Journal of the American Chemical Society Vol. 129; no. 14; pp. 4193 - 4206
Autores principales: Alfonso-Prieto, Mercedes, Borovik, Anton, Carpena, Xavier, Murshudov, Garib, Melik-Adamyan, William, Fita, Ignacio, Rovira, Carme, Loewen, Peter C.
Formato: Artículo
Publicado: American Chemical Society 4/11/2007
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Acceso en línea:Ver este registro en EBSCOhost
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        10.1021/ja063660y
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        atl: The Structures and Electronic Configuration of Compound I Intermediates of Helicobacter pylori and Penicillium vitale Catalases Determined by X-ray Crystallography and QM/MM Density Functional Theory Calculations.
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        au:
          Alfonso-Prieto, Mercedes
          Borovik, Anton
          Carpena, Xavier
          Murshudov, Garib
          Melik-Adamyan, William
          Fita, Ignacio
          Rovira, Carme
          Loewen, Peter C.
        affil:
          Centre especial de Recerca en Química Teòrica, Parc Científic de Barcelona, Josep Samitier 1-5, 08028 Barcelona, Spain
          Institute of Crystallography of Russian Academy of Sciences, Lenisky Prospekt 59, 119333 Moscow, Russia
          Institut de Biologia Molecular de Barcelona-CSIC, Parc Científic de Barcelona, Josep Samitier 1-5, 08028 Barcelona, Spain
          Institut de Recerca Biomèdica, Parc Científic de Barcelona, Josep Samitier 1-5, 08028 Barcelona, Spain
          Structural Biology Laboratory, Department of Chemistry, University of York, Heslington, York YO10 5YW, England
          Institució Catalana de Recerca i Estudis Avançats (ICREA), Passeig Lluís Companys 23, 08018 Barcelona, Spain
          Department of Microbiology, University of Manitoba, Winnipeg, Canada MB R3T 2N2
      su:
        Electronic structure
        Helicobacter pylori
        Penicillium
        Quantum theory
        X-ray crystallography
        Density functionals
        Physical vapor deposition
      sug:
        subj:
          Electronic structure
          Helicobacter pylori
          Penicillium
          Quantum theory
          X-ray crystallography
          Density functionals
          Physical vapor deposition
      ab: The structures of Helicobacter pylori (HPC) and Penicillium vitale (PVC) catalases, each with two subunits in the crystal asymmetric unit, oxidized with peroxoacetic acid are reported at 1.8 and 1.7 Å resolution, respectively. Despite the similar oxidation conditions employed, the iron-oxygen coordination length is 1.72 Å for PVC, close to what is expected for a FeO double bond, and 1.80 and 1.85 Å for HPC, suggestive of a FeO single bond. The structure and electronic configuration of the oxoferryl heme and immediate protein environment is investigated further by QM/MM density functional theory calculations. Four different active site electronic configurations are considered, PoṙFeO, PoṙFeO⋯HisH, PoṙFeOH and PorFeOH (a protein radical is assumed in the latter configuration). The electronic structure of the primary oxidized species, PoṙFeO, differs qualitatively between HPC and PVC with an A-like porphyrin radical delocalized on the porphyrin in HPC and a mixed A-like ‘fluctuating’ radical partially delocalized over the essential distal histidine, the porphyrin, and, to a lesser extent, the proximal tyrosine residue. This difference is rationalized in terms of HPC containing heme b and PVC containing heme d. It is concluded that compound I of PVC contains an oxoferryl PoṙFeO species with partial protonation of the distal histidine and compound I of HPC contains a hydroxoferryl PorFeOH with the second oxidation equivalent delocalized as a protein radical. The findings support the idea that there is a relation between radical migration to the protein and protonation of the oxoferryl bond in catalase.
      pubtype: Academic Journal
      doctype: Article
      src: R
    language: English
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