Structural Characterization of CO-Inhibited Mo-Nitrogenase by Combined Application of Nuclear Resonance Vibrational Spectroscopy, Extended X-ray Absorption Fine Structure, and Density Functional Theory: New Insights into the Effects of CO Binding and the Role of the Interstitial Atom

The properties of CO-inhibited Azotobader vinelandii (Av) Mo-nitrogenase (Nase) have been examined by the combined application of nuclear resonance vibrational spectroscopy (NRVS), extended X-ray absorption fine structure (EXAFS), and density functional theory (DFT). Dramatic changes in the NRVS are...

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Publicado en:Journal of the American Chemical Society Vol. 136; no. 45; pp. 15942 - 15955
Autores principales: Scott, Aubrey D., Yan, Lifen, George, Simon J., Wang, Honexin, Cramer, Stephen P., Pelmenschikov, Vladimir, Guo, Yisong, Dapper, Christie H., Newton, William E., Yoda, Yoshitaka, Tanaka, Yoshihito
Formato: Artículo
Publicado: American Chemical Society 11/12/2014
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Acceso en línea:Ver este registro en EBSCOhost
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      dt: 11/12/2014
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      pub: American Chemical Society
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        10.1021/ja505720m
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        atl: Structural Characterization of CO-Inhibited Mo-Nitrogenase by Combined Application of Nuclear Resonance Vibrational Spectroscopy, Extended X-ray Absorption Fine Structure, and Density Functional Theory: New Insights into the Effects of CO Binding and the Role of the Interstitial Atom
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          Scott, Aubrey D.
          Yan, Lifen
          George, Simon J.
          Wang, Honexin
          Cramer, Stephen P.
          Pelmenschikov, Vladimir
          Guo, Yisong
          Dapper, Christie H.
          Newton, William E.
          Yoda, Yoshitaka
          Tanaka, Yoshihito
        affil:
          Department of Chemistry, University of California, Davis, California 95616, United States
          Physical Biosciences Division, Lawrence Berkeley National Laboratory, Berkeley, California 94720, United States
          Institut für Chemie, Technische Universität Berlin, 10623 Berlin, Germany
          Department of Chemistry, Carnegie Mellon University, Pittsburgh, Pennsylvania 15213, United States
          Department of Biochemistry, Virginia Polytechnic Institute & State University, Blacksburg, Virginia 24061, United States
          Research and Utilization Division, SPring-8/JASRI, 1-1-1 Kouto, Sayo, Hyogo 679-5198, Japan
          SR Materials Science Instrumentation Unit, RIKEN SPring-8 Center, 1-1-1 Kouto, Sayo, Hyogo 679-5148, Japan
      su:
        Nitrogenases
        Nuclear resonance reactions
        Vibrational spectra
        X-ray absorption
        Density functional theory
        Interstitial defects
        Atoms
      sug:
        subj:
          Nitrogenases
          Nuclear resonance reactions
          Vibrational spectra
          X-ray absorption
          Density functional theory
          Interstitial defects
          Atoms
      ab: The properties of CO-inhibited Azotobader vinelandii (Av) Mo-nitrogenase (Nase) have been examined by the combined application of nuclear resonance vibrational spectroscopy (NRVS), extended X-ray absorption fine structure (EXAFS), and density functional theory (DFT). Dramatic changes in the NRVS are seen under high-CO conditions, especially in a 188 cm mode associated with symmetric breathing of the central cage of the FeMo-cofactor. Similar changes are reproduced with the α-H195Q_ N2ase variant. In the frequency region above 450 cm, additional features are seen that are assigned to Fe-CO bending and stretching modes (confirmed by CO isotope shifts). The EXAFS for wild-type Nase shows evidence for a significant cluster distortion under high-CO conditions, most dramatically in the splitting of the interaction between Mo and the shell of Fe atoms originally at 5.08 A in the resting enzyme. A DFT model with both a terminal - CO and a partially reduced - CHO ligand bound to adjacent Fe sites is consistent with both earlier FT-IR experiments, and the present EXAFS and NRVS observations for the wild-type enzyme. Another DFT model with two terminal CO ligands on the adjacent Fe atoms yields Fe-CO bands consistent with the a-H195Q variant NRVS. The calculations also shed light on the vibrational "shake" modes of the interstitial atom inside the central cage, and their interaction with the Fe-CO modes. Implications for the CO and N reactivity of Nase are discussed.
      pubtype: Academic Journal
      doctype: Article
      src: R
    language: English
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          year: 2014
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